Behavior
The groups consist of up to four adult males and many adult females; typically yielding a 1:2 ratio of males to females. Young of varying ages are also incorporated in the group. The number of monkeys in a group can range from thirty to fifty individuals. The species is a very social animal, and can often be observed playing and grooming during the rest periods between meals. Unlike females, in a group, males actually maintain close bonds, acting together in defense of their group and even in grooming each other.
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—Elaine Heffner (20th century)