Awards
- 1978 Foreign Honorary Member of American Academy of Arts and Sciences
- 1980 Fellow of the Royal Society of London
- 1982 Newcomb Cleveland Prize of the American Association for the Advancement of Science
- 1988 Darwin Medal of the Royal Society of London
- 1989 Scientific Medal of the Linnean Society
- 1991 Frink Medal of Zoological Society of London
- 1992/3 Wander Prize of the University of Bern
- 1993 Crafoord Prize of the Royal Swedish Academy of Sciences
- 1993 Kyoto Prize of the Inamori Foundation
- 1995 Fyssen Prize of the Fyssen Foundation
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