Saturation
Although not strictly necessary for a neutral theory, many stochastic models of biodiversity assume a fixed, finite community size. There are unavoidable physical constraints on the total number of individuals that can be packed into a given space (although space per se isn't necessarily a resource, it is often a useful surrogate variable for a limiting resource that is distributed over the landscape; examples would include sunlight or hosts, in the case of parasites).
If a wide range of species is considered (say, giant sequoia trees and duckweed, two species that have very different saturation densities), then the assumption of constant community size might not be very good, because density would be higher if the smaller species were monodominant.
However, because the Unified Theory refers only to communities of trophically similar, competing species, it is unlikely that population density will vary too widely from one place to another.
Hubbell considers the fact that population densities are constant and interprets it as a general principle: large landscapes are always biotically saturated with individuals. Hubbell thus treats communities as being of a fixed number of individuals, usually denoted by J.
Exceptions to the saturation principle include disturbed ecosystems such as the Serengeti, where saplings are trampled by elephants and Blue wildebeests; or gardens, where certain species are systematically removed.
Read more about this topic: Unified Neutral Theory Of Biodiversity
Famous quotes containing the word saturation:
“In the twentieth century, death terrifies men less than the absence of real life. All these dead, mechanized, specialized actions, stealing a little bit of life a thousand times a day until the mind and body are exhausted, until that death which is not the end of life but the final saturation with absence.”
—Raoul Vaneigem (b. 1934)