Plot
The film begins with a summary of how the Allies are struggling to stop U-boats from sinking their freighters. The scene transfers to U-571, which torpedoes and sinks a freighter. The crew is happy with the kill, but seconds later the sonar man reports having detected high speed screws approaching. The Captain sights a destroyer, and orders the U-571 to dive. The destroyer drops depth charges, snapping a fuel line, which catches fire and kills the entire engineering crew. Because of sustained damage, U-571 is forced to resurface. The Captain learns that the batteries are almost depleted, both diesel engines are inoperable, and the engineering crew is dead. An Enigma-encoded SOS is sent to Berlin for aid.
Meanwhile, the crew of the United States Navy submarine S-33 are celebrating ENS Larson's wedding and 48 hours leave. During the party, LT Andrew Tyler enters looking solemn, having been denied his first command. After complaining to LCDR Dahlgren, he is rebuffed and upset to learn that Dahlgren withheld his recommendation and voted against his receiving the promotion. Military Police suddenly arrive, announcing a secret mission that ends shore leave. The men arrive to find their boat, the S-33, modified to resemble a German U-boat. The crewman must wear German uniforms. Hirsch, a naval Intelligence officer who is fluent in German, orders Tyler to locate Radioman Wentz, who is fluent in German due to his immigrant parents. At the same time, a Marine named Coonan arrives in a convoy loaded with high explosives. After the S-33 sails, Hirsch explains that the Allies intercepted the disabled U-571's SOS. They will masquerade as the resupply ship for U-571, board it, capture the Enigma coding device and scuttle (sink) the U-571. Tyler is skeptical but goes along. Wentz asks Tyler to keep his German heritage a secret. The S-33 leaves Portsmouth Naval Shipyard in Portsmouth, New Hampshire.
Back on U-571, attempted repairs fail and the Captain is alerted that survivors from the merchant ship he sank have been spotted in a lifeboat, asking for asylum. He orders his men to shoot them, as their orders are to spare no survivors. His men reluctantly comply.
During a storm, the S-33 comes across U-571 and sends the boarding party over, led by Coonan. Hirsch temporarily freezes and Wentz is forced to speak German in front of his friends in order for the group to retain their cover until their rafts are tied up. At first, the Germans openly welcome the S-33, but when a midshipman looks through his binoculars, he immediately identifies the weapons as American. He alerts the rest of the Germans, and they exchange fire with the S-33 crewmen. The Americans are able to take the boat by force, capture the Enigma and begin rounding up the prisoners, including the Captain. As the prisoners are transferred between ships and the scuttling charges are laid, Dahlgren, standing on the upper deck of the S-33, notices the sound of a detonation. He turns around in horror, to see a streak coming towards the S-33 underwater. The ship is torpedoed and sunk by the real German resupply sub. C.O. Dahlgren, wounded in the water, orders his men on the captured U-boat to submerge and save themselves. (This scene was based on an actual incident in World War II: Commander Howard W. Gilmore, USN, after being seriously wounded in an encounter with a Japanese destroyer, ordered his men to abandon him on deck and submerge to save the ship and crew. Gilmore was posthumously awarded the Medal of Honor for his sacrifice. The movie quotes Gilmore's last recorded words: "Take her down.") Coonan, Larson and many others are lost, forcing Tyler to take command and dive the captured U-boat. There is a desperate scramble to disarm the charges they placed. With only Hirsch and Wentz able to read German, they struggle with interpreting the controls and then fire a salvo of torpedoes, destroying the enemy U-Boat, draining the last of the sub's batteries and spending all but the last torpedo, loaded in the malfunctioning aft torpedo tube. Surfacing, Tyler and his men search for survivors and find two: Eddie the cook from the S-33 and a German sailor claiming to be an electrician, but who is actually the Captain of U-571.
After repairing one of the diesel engines, thus restoring power and propulsion, Tyler decides to take the disabled submarine to England. Some of the men disagree with Tyler's decision and Tyler replies with, "I don't know", to their questions. Chief Klough privately rebukes Tyler, saying, "A Captain always knows what to do, whether he does or not." He also rebukes Mazzola in front of others for openly disagreeing with Tyler. They spot an aircraft, and Mazzola - against Tyler's orders - tries to convince Rabbit to fire on the plane with the deck gun, which appears to be coming in for an attack, but is only scouting for a German destroyer named the Anschluss. Tyler punches Mazzola for ignoring the Chain of Command and his stupidity. The captured German Captain breaks free, attacks Tank and kills Mazzola before being subdued. Unaware that the U-571 has been commandeered by Americans, the Anschluss sends over a small contingent to meet and greet their German comrades. Right before boarders arrive, Tyler has Rabbit fire a shot from the deck gun right into the ship's radio tower, preventing the destroyer from reporting that the Enigma code has been compromised, and dives underneath her. The destroyer begins to drop depth charges to try to sink U-571.
Tyler plans to trick the destroyer into stopping by ejecting debris and Mazzola's corpse out of an empty torpedo tube, faking their own destruction. The younger men balk at using their crewman's body this way, but Tyler states that Mazzola is saving their lives. U-571 will then surface and hit the ship with their last torpedo.
The German destroyer continues dropping depth charges. U-571, hiding at great depth below 200 meters, is damaged by the high water pressure. In preventing the submarine from sinking, control of the main ballast tanks are lost and the ship ascends uncontrollably. Tyler orders Trigger to submerse himself in the bilge underwater to repressurize the torpedo tubes. During the ascent, the German prisoner tries to warn the destroyer in Morse Code that they're not dead yet. Wentz translates the message in German, "I am U-571. Destroy me!" Enraged, Hirsch grabs a large wrench and kills the prisoner. Trigger manages to close the air valve for the tubes, but a second leak and valve are unexpectedly revealed, both behind a wall of pipes; Trigger's arm can't reach, and his air hose is too short. When Tank reports this, Tyler rushes to the engine room himself. Tyler tells him that they need him to do this and orders him to get the job done. U-571 surfaces without a torpedo to fire. The destroyer fires on the ship, which runs using its diesel engine, but takes heavy damage from the destroyer's deck guns and starts to flood. Trigger leaves behind the air hose and closes the second valve, but the damage causes pipes to collapse, trapping his leg, and he drowns. The second the pressure is available, Tyler orders Tank to fire the final torpedo. The German ship is destroyed; in front of everyone, Chief Klough tells Tyler that if he ever needs a Chief, he would gladly go to sea with him anytime. Tank reports Trigger's death while carrying out his order, but U-571 has taken severe damage and will not stay afloat for long. The crew abandons ship with the Enigma in tow, and watches it sink while mourning their lost crewmates and for the German sub, which ironically, saved their lives. Floating aboard an inflatable lifeboat, they are eventually spotted by a US Navy sea-plane.
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