Anatomy
The olfactory tubercle is generally located at the basal forebrain of the animal within the medial temporal lobe. Specifically parts of the tubercle are included in the olfactory cortex and nested between the optic chiasm and olfactory tract and ventral to the nucleus accumbens. The olfactory tubercle consists of three layers, a molecular layer (layer I), the dense cell layer (layer II) and the multiform layer (layer III) (Millhouse & Heimer 1984). Other than the islands of calleja which are characteristic of the tubercle (Calleja 1893), it is also noted for the being innervated by dopaminergic neurons from the ventral tegmental area. The olfactory tubercle also consist of heterogeneous elements such as medial forebrain bundle and has a ventral extension of the striatal complex. During the 1970s the tubercle was found to contain a striatal component which is filled GABAergic medium spiny neurons. The GABAergic project to the ventral pallidum and receive glutamatergic inputs from cortical regions and dopaminergic inputs from the ventral tegmental area (Millhouse 1987; Meyer 1989).
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