Education
Graduated from Iroquois Central High School, Elma, New York, in 1978. Went on to earn two bachelor of arts degrees in biology and psychology from the University of Rochester, New York in 1982. At Pennsylvania State University he earned a master of science degree in physiology in 1985, and a doctor of philosophy degree in biology (physiology) from the University of North Texas in 1989. He completed a post-doctoral fellowship at the University of Texas Southwestern Medical Center in 1992.
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